1.8.3.3 C124A reduced activity compared to wild type enzyme 1.8.3.3 C15A reduced activity compared to wild type enzyme 1.8.3.3 C15A/C124A reduced activity compared to wild type enzyme 1.8.3.3 C15A/C74A/C85A/C124A reduced activity compared to wild type enzyme 1.8.3.3 C74A/C85A reduced activity compared to wild type enzyme