1.3.5.6 FAD tightly bound to zeta-carotene desaturase CrtQb 1.3.5.6 additional information addition of the dinucleotides FAD, NADP+ or NAD+ alone or the latter in combination with ATP results in a negligible increase in zeta-carotene desaturase activity 1.3.5.6 quinone - 1.3.5.6 quinone the expression of zeta-carotene desaturase paralleles plastid terminal oxidase (PTOX, quinol:oxygen oxidoreductase) and zeta-carotene desaturase. The three genes are expressed when carotinoid biosynthesis is enhanced during fruit ripening. Tomato ghost mutant with impaired plastid terminal oxidase accumulate phytoene in leaves and fruits