1.13.11.4 gentisate 1,2-dioxygenase gentisate degradation I - - PWY-6223 3.7.1.20 3-fumarylpyruvate hydrolase gentisate degradation I - - PWY-6223 5.2.1.4 maleylpyruvate isomerase gentisate degradation I - - PWY-6223