1.1.1.330 very-long-chain 3-oxoacyl-CoA reductase (4Z,7Z,10Z,13Z,16Z)-docosa-4,7,10,13,16-pentaenoate biosynthesis II (4-desaturase) 1.3.1.93 very-long-chain enoyl-CoA reductase (4Z,7Z,10Z,13Z,16Z)-docosa-4,7,10,13,16-pentaenoate biosynthesis II (4-desaturase) 4.2.1.134 very-long-chain (3R)-3-hydroxyacyl-CoA dehydratase (4Z,7Z,10Z,13Z,16Z)-docosa-4,7,10,13,16-pentaenoate biosynthesis II (4-desaturase)