1.2.1.74 abieta-7,13-dien-18-al dehydrogenase abietic acid biosynthesis - 1.14.14.144 abieta-7,13-diene hydroxylase abietic acid biosynthesis - 4.2.3.18 abieta-7,13-diene synthase abietic acid biosynthesis - 5.5.1.12 copalyl diphosphate synthase abietic acid biosynthesis -