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Catalyses a step in the 3-hydroxypropanoate/4-hydroxybutanoate cycle, an autotrophic CO2 fixation pathway found in some thermoacidophilic archaea . The enzyme from Sulfolobus tokodaii does not act on either NADH or crotonyl-CoA . Different from EC 18.104.22.168, which acts only on enoyl-CoA derivatives of carbon chain length 4 to 16. Contains Zn2+.